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You are here: Home / Aquatic Species / Species Profiles / Clubbed Tunicate
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Species Profiles

Clubbed Tunicate

Clubbed tunicate - Washington Department of Fish & WildlifeScientific name: Styela clava Herdman, 1881

Common names: Clubbed tunicate, stalked sea squirt, solitary sea squirt, leathery sea squirt, Pacific rough sea squirt, Asian sea squirt

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Native To: Eastern Asia (Clarke and Therriault 2007)

Date of U.S. Introduction: First observed on the Pacific coast in the 1930s (Clarke and Therriault 2007)

Images: Google

Means of Introduction: Probably introduced accidentally on the hulls of ships or with imported oysters (Clarke and Therriault 2007)

Impact: Competes with and displaces native species (Clarke and Therriault 2007)

Current U.S. Distribution: Coastal regions of New England and the Western U.S.

Management Plans: Aquatic Species

Selected Internet Resources:

Federal Government

Styela clava
Integrated Taxonomic Information System.
Taxonomy

Styela clava Article Citation Search - AGRICOLA Database
USDA. National Agricultural Library.
Research

Styela clava - Rhode Island Marine & Estuarine Invasive Species Site
DOC. NOAA. National Estuarine Research Reserve System. Narragansett Bay Research Reserve. Coastal Training Program.
Photographs; Introduction History; Impacts; Life Cycle; Habitat; Distribution

Marine Invasive Species: Styela clava (PDF | 84 KB)
DOI. NPS. Natural Resource Program Center.
Photographs; Impacts; Habitat; Distribution; Dispersion; Controls

Styela clava - National Exotic Marine and Estuarine Species Information System, Chesapeake Bay Introduced Species Database
Smithsonian Institution. Smithsonian Environmental Research Center.
Taxonomy; Identification/Description; Photographs; Introduction History; Habitat; Distribution; Dispersion

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State Government

Styela clava (Clubbed Tunicate)
Washington Department of Fish and Wildlife.
Identification/Description; Photographs; Introduction History; Impacts

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University/Academic

Invasive Tunicate Identification Card: Colonial Tunicate and Club Tunicate (PDF | 1.6 MB)
University of Washington. Washington Sea Grant Program.
Identification/Description; Photographs; Habitat

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International

Styela clava - Invasive Species Compendium
Centre for Agricultural Bioscience International (CABI).
Taxonomy; Identification/Description; Photographs; Impacts; Life Cycle; Habitat; Distribution; Controls

Biological Synopsis of the Invasive Tunicate Styela clava (Herdman 1881) (2007; PDF | 1.4 MB)
Fisheries and Oceans Canada.
Taxonmy; Identification/Description; Introduction History; Impacts; Life Cycle; Habitat; Distribution; Dispersion

Styela clava - Delivering Alien Invasive Species Inventories for Europe (Jan 2008; PDF | 163 KB)
European Commission. Sixth Framework Programme.
Taxonomy; Identification/Description; Photographs; Introduction History; Impacts; Life Cycle; Habitat; Distribution; Dispersion; Controls

Sea Squirt (Clubbed Tunicate)
New Zealand Ministry of Agriculture and Forestry. Biosecurity New Zealand.
Identification/Description; Photographs; Impacts; Habitat; Distribution; Dispersion

Invasive Sea Squirt (Styela clava) Fact Sheet
National Institute of Water and Atmospheric Research (New Zealand).
Identification/Description; Photographs; Impacts; Habitat; Distribution; Dispersion; Controls

Leathery Sea Squirt (Dec 2006; PDF | 110 KB)
Swedish Environmental Protection Agency. Främmande Arter i Svenska Hav (Alien Species in Swedish Seas).
Identification/Description; Photographs; Introduction History; Impacts; Habitat; Distribution

Non-Native Species: Styela clava
Joint Nature Conservation Committee (United Kingdom).
Taxonomy; Introduction History; Impacts; Distribution; Dispersion

A sea squirt - Styela clava
Marine Biological Association (United Kingdom). Marine Life Information Network for Britain and Ireland.
Taxonomy; Identification/Description; Photographs; Introduction History; Habitat; Distribution; Dispersion

Styela clava, Leathery sea squirt - Invasive Alien Species in Northern Ireland
National Museums Northern Ireland (United Kingdom).
Identification/Description; Impacts; Life Cycle; Distribution; Dispersion; Controls

Styela clava (tunicate) - ISSG Global Invasive Species Database
World Conservation Union. Invasive Species Specialist Group.
Identification/Description; Impacts; Life Cycle; Habitat; Distribution; Dispersion; Controls

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Organizations

Styela clava - Marine Organisms Database
Marine Biological Laboratory (Massachusetts).
Taxonomy; Photographs; Introduction History; Impacts; Distribution

Club tunicate
Northeast Aquatic Nuisance Species Panel.
Identification/Description; Photographs; Introduction History; Impacts; Habitat; Distribution

Styela clava - Guide to Marine Invaders in the Gulf of Maine (PDF | 311 KB)
Salem Sound Coastwatch (Massachusetts).
Identification/Description; Photographs; Illustrations; Introduction History; Habitat; Dispersion

Styela clava Herdman, 1881 - Guide to the Exotic Species of San Francisco Bay
San Francisco Estuary Institute.
Taxonomy; Identifcation/Description; Photographs; Introduction History; Impacts; Life Cycle; Habitat; Distribution

Invasive Tunicates in Washington State: Styela clava or Club tunicate
Washington Scuba Alliance.
Identification/Description; Photographs; Impacts; Habitat; Distribution

Citations

Clarke, C.L. and T.W. Therriault. 2007. Biological Synopsis of the Invasive Tunicate Styela clava (Herdman 1881) (PDF | 1.38 MB) Pacific Biological Station, Fisheries and Oceans Canada.

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Last Modified: Mar 24, 2014
 
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